The head of the family is the senior representative of the armigerous families of Willis of Bletchley and Fleming of Stoneham, and is a direct descendant in the male line of Dr Thomas Willis.
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| 1670-1675 | Dr Thomas Willis (1621-1675)
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| 1675-1699 | Thomas Willis (1658-1699)
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| 1699-1760 | Browne Willis (1682-1760)
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| 1760-1762 | Thomas Willis (1738-1762)
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| 1770-1802 | John Willis, subsequently known as John Fleming (1749-1802)
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| 1813-1844 | John Fleming Barton Willis, subsequently known as John Willis Fleming (1781-1844)
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| 1844-1872 | John Brown Willis Fleming (1815-1872)
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| 1893-1949 | John Edward Arthur Willis Fleming (1871-1949)
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| 1949-1967 | John Brown Phillimore Willis Fleming (1895-1967)
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| 1967-2006 | Richard Hugh Willis Fleming (1921-2006)
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| From 2006 | John Valentine Willis Fleming (b.1925)
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